Article © D & R Lalkaka, uploaded January 01, 2003.

Dinyar Lalkaka introduces this extravagant fish - a species that burns holes in wallets the world over. In the UK, one grand (a thousand pounds) was, at one time, the exorbitant asking price for one of these inky beauties. The term, appropriately, gives rise to their fishkeeping slang name of Synodontis grandulosus! I'll leave it with Dinyar now to further discuss this lacustrine gem...

Few catfish have acquired more mystique than the majestic Synodontis granulosus, one of six described Synodontis species endemic to Lake Tanganyika. In part, this mystique reflects the princely price the fish commands. When available, it is often the most expensive fish on vendors' sales lists. Whereas the prices of nearly all once-expensive catfish have deflated rapidly over the last decade, the going rates for S. granulosus have come down only slightly, dropping from around US $500 as recently as early 2000 to about $300 today (and about 300 in Europe). If you are going to pay several hundred dollars for a fish, you'd better earn lots of bragging points with your fish buddies. And have a good explanation for your spouse!

A second reason for the high price and mystique of S. granulosus is that the fish is relatively uncommon in the wild. Unlike many Lake Tanganyika Synodontis, it is a loner, and thus impossible to catch in large numbers. Moreover, it is found in deep water (about 50 meters) at some distance from the shore. Until a few years ago, this was beyond the limits of existing diving technology and the fish were mostly caught in gill nets, a low yield method which restricted the volume of exports. As a result, only 20-30 fish were exported from the Lake each year. Better diving technology has changed this situation, and it appears that many hundred fish are now shipped out of the Lake annually. Whether this is a sustainable level remains to be seen. There is already some evidence that the rocky habitat of S. granulosus is under growing ecological pressure.

Synodontis granulosus

The third and best reason for the extraordinary mystique of S. granulosus is that it is truly an exceptionally elegant and intelligent fish that quickly becomes the cynosure of any Tanganyika tank. The body is brown-black, shot with a purple sheen and highlighted with bright white barbels and bright white edging on the posterior borders of its fins. Although not evident at first sight, close inspection reveals that the skin of the whole body, including the adipose fin, is covered with small, closely set granular papilla (hence the name "granulosus"). Most specimens are imported at a size of 12-18 cm, but mature adults can reach 25 cm SL.

S. granulosus quickly learns who the "Food Guy" is and will swim up to be fed by hand. Every time I put my hand into its tank, my "granny" comes and nibbles at it! (It tickles, but unlike the nipping of S. polli, which sometimes do the same thing, doesn't hurt. My three year old daughter asks, "Daddy, why does this fish keep licking you?") Many aficionados of Tanganyika fish keep their S. granulosus with herbivorous fish like Tropheus, and are still able to supplement their diet with meaty fare by hand feeding them. They have a keen sense of smell and are usually the first to get at food. When I feed my Tanganyika tank, my granny races across the surface, upside down in classic Synodontis-style, vacuuming up everything she can. Most specimens are voracious eaters and can easily become overfed in community tanks. If S. granulosus has a fault, it is gluttony.

In smaller tanks, S. granulosus tends to be both reclusive and aggressive towards other Synodontis, especially conspecifics. Females tend to be a bit larger and more aggressive than the males. If you are lucky enough to keep several S. granulosus, it's best to keep four or more in order to spread out aggression. Some specimens are bolder than others, but almost all will become the life and soul of the party if kept in a tank of 125 gallons (about 500 liters) or more, with lots of rockwork, ample open swimming space above the rocks, and the company of schools of other Lake Tanganyika Synodontis species.

Adult S. granulosus are easy to identify, unlike the other five Synodontis species endemic to Lake Tanganyika. Moreover, the taxonomic status of S. granulosus is so clear that in the 103 years since it was first described by George Boulenger in 1900, not a single synonym has been introduced. By comparison, taxonomic confusion still swirls around the identities of other Lake Tanganyika Synodontis such as S. petricola, Matthes, 1959, which clearly needs to be divided into multiple separate species, and to a lesser extent, S. tanganaicae and S. dhonti.

Venn diagram Lake Tanganyika Synodontis spp.

The distinctiveness of S. granulosus is easy to see in the Venn diagram at the right. Some Tanganyika Synodontis are granulose, like S. granulosus (though to a lesser extent), but all except S. granulosus are distinctly spotted.

While the diagram accurately describes Tanganyika Synodontis as typically seen in the aquarium, in fact, it's not quite that clear cut, because juvenile and sub-adult S. granulosus also have spots (not to mention the fact that all six endemic species have white fin edging to some degree). Indeed, very young granulosus have a color pattern that, to the untrained eye, could easily be mistaken for S. multipunctatus! In the picture below of a sub-adult specimen, faint spots can still be seen. Moreover, fully mature S. dhonti and S. tanganaicae lose their spots.

Synodontis granulosus

This illustrates a general fact about the entire Lake Tanganyika Synodontis species flock: fry and juveniles of all endemic species look very similar: the distinctive characteristics of each species only become fully apparent as the fish reach adulthood.

Could this be the reason that very young wild-caught S. granulosus have apparently never entered the trade? In any event, little is known about S. granulosus' spawning habits, and the fish has never been spawned naturally by aquarists. While it has been bred by hormone injection in it While it has been bred by hormone injection in the Czech Republic, it is reliably reported that these artificially spawned fish are the result of hybridization by unscrupulous breeders and lose their resemblance to true S. granulosus as they mature. Given the limited numbers of S. granulosus in the wild, its apparently threatened habitat and its extraordinary elegance and charm, it is only to be hoped that future captive breeding efforts can lead to the emergence of a healthy and sustainable tank-raised population spawned without recourse to artificial hybridization. Planet Earth would shine less brightly were this elegant fish to disappear.

Special thanks to Greg Pierson for his major contributions to this article.

Copyright information for the images used in this article can be found on the species' full Cat-eLog page.

Cat-eLog Data Sheet
Scientific Name	Synodontis granulosus  Boulenger, 1900
Common Names	White Seam Syno Deep Lake Syno, Granny, Hvidsømmet Skægmalle (Denmark), Leuchtbaken-Fiederbartwels (Germany)
Type Locality	Northern end of Lake Tanganyika.
Synonym(s)	Synodontis granulosa
Pronunciation	sin oh don tiss - gran you LOW suss
Etymology	According to Cuvier, Synodontis is an "ancient name for an undetermined fish from the Nile". It is not derived, as often reported, from syn-, together and odous, tooth, presumed etymology of the lizardfish genus Synodus and refers to the closely-spaced lower jaw teeth of both genera.
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Species Information
Size	270mm or 10.6" SL. Find near, nearer or same sized spp.
Identification	All species in the genus Synodontis have a hardened head cap that has attached a process (humeral process) which is situated behind the gill opening and pointed towards the posterior. The dorsal fin and pectoral fins have a hardened first ray which is serrated. Caudal fin is always forked. There is one pair of maxillary barbels, sometimes having membranes and occasionally branched. The two pairs of mandibular barbels are often branched and can have nodes attached. The cone-shaped teeth in the upper jaw are short. S-shaped and movable in the lower jaw. These fish produce audible sounds when disturbed rubbing the base of the pectoral spine against the pectoral girdle. At least 1/3 to 1/2 of all fins have a snow white edging. Long white barbles. Inky sheen to brownish body which bears a spotted pattern in younger fish. The contrast of this spotting diminishes with age.
Sexing	Both sexes have identical colouration. Females grow slightly larger and tend to be plumper than males. Males have a 4-5 mm genital papilla near the anus.
Habitat Information
Distribution	Africa: Lake Tanganyika. African Waters, Western Rift Valley Lakes, Tanganyika (click on these areas to find other species found there) Login to view the map.
IUCN Red List Category	Least Concern , range map and more is available on the IUCN species page. Last assessed 2006.
pH	7.8 - 8.4
Temperature	24.0-28.0°C or 75.2-82.4°F (Show species within this range)
Husbandry Information
Feeding	Voracious eater that will accept virtually all aquarium food, but prefers meaty foods such as shrimp and clam. Apparently feeds on small molluscs in the wild. User data.
Furniture	A network of caves formed by piling up rocks, with ample swimming space above. Vallisneria and / or hornwort is optional.
Compatibility	Best kept in a tank of 125 gallons (500 liters) or more. May become reclusive and aggressive in tanks smaller than 70 gallons (250 liters). May jump out of tank in first few days after initial introduction.
Suggested Tankmates	Tanganyikan Synodontis, robust Tanganyikan cichlids and Tanganyika eels.
Breeding	Breeding has been reported by a UK based wholesaler, but no details have been made available. Has also been bred by hormone injection in Eastern Europe.
Breeding Reports	There is no breeding report.
Further Information
Reference	Annals and Magazine of Natural History (Series 7) v. 6 (no. 35), pp 480.
Registered Keepers	There are 41 registered keepers, view all "my cats" data.
Wishlists	Love this species? Click the heart to add it to your wish list. There are 9 wishes to keep this species, see who wants what.
Spotters	Spotted this species somewhere? Click the binoculars! There are 19 records of this fish being seen, view them all.
Forum BBCode
	Search for S. granulosus
	Look up S. granulosus on AquaticRepublic.com
	Look up S. granulosus on Fishbase
	Look up S. granulosus on Encyclopedia of Life
	Look up S. granulosus on Global Biodiversity Information Facility
	LFS label creator ARN ref:1.13.29.416
Last Update	2020 Sep 19 01:40 (species record created: 2003 Jan 01 11:22)

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